Earth Sciences / Population Biology / Stratification / Biological Sciences / Environmental Sciences / Computer Model / Density dependence / Chironomidae / Food Availability / Spatial Distribution / Temperature Dependence / Three Dimensional / Water Level / Development Time / Qualitative Modeling / Computer Model / Density dependence / Chironomidae / Food Availability / Spatial Distribution / Temperature Dependence / Three Dimensional / Water Level / Development Time / Qualitative Modeling